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Centipedes (from Latin prefix centi-, "hundred", and pes, pedis, "foot") are arthropods belonging to the class Chilopoda of the subphylum Myriapoda. They are elongated metameric animals with one pair of legs per body segment. Despite the name, centipedes can have a varying number of legs from under 20 to over 300. Centipedes have an odd number of pairs of legs, e.g. 15 or 17 pairs of legs (30 or 34 legs) but never 16 pairs (32 legs). A key trait uniting this group is a pair of venom claws or "forcipules" formed from a modified first appendage. Centipedes are a predominantly carnivorous taxon.:168 Centipedes normally have a drab coloration combining shades of brown and red. Cavernicolous (cave-dwelling) and subterranean species may lack pigmentation and many tropical scolopendromorphs have bright aposematic colours. Size can range from a few millimetres in the smaller lithobiomorphs and geophilomorphs to about 30 cm (12 in) in the largest scolopendromorphs. Centipedes can be found in a wide variety of environments. Worldwide there are estimated to be 8,000 species of centipede, of which 3,000 have been described. Centipedes have a wide geographical range, reaching beyond the Arctic Circle. Centipedes are found in an array of terrestrial habitats from tropical rainforests to deserts. Within these habitats centipedes require a moist micro-habitat because they lack the waxy cuticle of insects and arachnids, and so lose water rapidly through the skin. Accordingly, they are found in soil and leaf litter, under stones and dead wood, and inside logs. Centipedes are among the largest terrestrial invertebrate predators and often contribute significantly to the invertebrate predatory biomass in terrestrial ecosystems.