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Ocypodinae
Most ghost crabs have pale-colored bodies that blend in well with the sand, though they are capable of gradually changing body coloration to match their environments and the time of day. Some species are brightly colored, such as Ocypode gaudichaudii and Ocypode ryderi. Ghost crabs have elongated and swollen eyestalks with very large corneas on the bottom half. Their carapaces are deep and box-like, squarish when viewed from the top with straight or slightly curving sides. The regions of the carapace are usually not clearly defined. The "whip" of their antennules (antennular flagella) are small or rudimentary. They fold back into the body diagonally or almost vertically. The plate between them (the interantennular septum) is broad. The third pair of mouth appendages (maxillipeds) completely cover the mouth opening. A small orifice with edges thickly fringed with hair is found between the bases of the second and third pairs of walking legs.
Ghost crabs dominate sandy shores in tropical and subtropical regions of the world, replacing the sandhoppers that predominate in cooler areas. Three species are found in the Atlantic Ocean and the Mediterranean Sea, and two occur in the eastern Pacific coast of the Americas. The rest of the species are found in the western Pacific and the Indian Ocean to the tip of southern Africa.
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