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These are golden-brown sori (collection of sporangia/fruiting bodies) on the underside of a fern frond.
Grows in the cracks of a shale wall across from a waterfall, a high humidity area at Watkins Glen State Park.
The presence and morphology of the sori helps to identify or confirm the fern species.
Lat: 42.37, Long: -76.89
Spotted on Jun 1, 2019
Submitted on Jul 4, 2019