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These are golden-brown sori (collection of sporangia/fruiting bodies) on the underside of a fern frond.
Grows in the cracks of a shale wall across from a waterfall, a high humidity area at Watkins Glen State Park.
The presence and morphology of the sori helps to identify or confirm the fern species.
Spotted on Jun 1, 2019
Submitted on Jul 4, 2019